Delhey plumage lightness benchmark

Environmental covariates for trait imputation on 5809 passerine species (Delhey et al. 2019).

Traits: lightness_male, lightness_female

Covariates: annual_mean_temperature, annual_precipitation, percent_tree_cover, mean_temperature_of_warmest_quarter, precipitation_of_warmest_quarter, midLatitude

Test-set RMSE

Missing: 20%

Trait Mean imputation BM baseline pigauto (no covariates) pigauto + covariates
lightness_male 1.0062 0.5961 0.5961 0.5956
lightness_female 1.0139 0.6259 0.6239 0.6275

Missing: 40%

Trait Mean imputation BM baseline pigauto (no covariates) pigauto + covariates
lightness_male 1.0239 0.6473 0.6473 0.6473
lightness_female 0.9911 0.7063 0.7073 0.7063

Missing: 60%

Trait Mean imputation BM baseline pigauto (no covariates) pigauto + covariates
lightness_male 0.9807 0.6943 0.6943 0.6943
lightness_female 0.9757 0.7273 0.7273 0.7262

Test-set Pearson r

Missing: 20%

Trait Mean imputation BM baseline pigauto (no covariates) pigauto + covariates
lightness_male 0.8067 0.8067 0.8064
lightness_female 0.7890 0.7912 0.7889

Missing: 40%

Trait Mean imputation BM baseline pigauto (no covariates) pigauto + covariates
lightness_male 0.7759 0.7759 0.7759
lightness_female 0.7120 0.7116 0.7120

Missing: 60%

Trait Mean imputation BM baseline pigauto (no covariates) pigauto + covariates
lightness_male 0.7151 0.7151 0.7151
lightness_female 0.6842 0.6842 0.6840

Covariate lift summary

RMSE ratio of pigauto + covariates relative to pigauto (no covariates). Values < 1 indicate covariates help.

Missing %TraitRMSE (no cov)RMSE (with cov)Ratio
20% lightness_female 0.6239 0.6275 1.006
20% lightness_male 0.5961 0.5956 0.999
40% lightness_female 0.7073 0.7063 0.999
40% lightness_male 0.6473 0.6473 1.000
60% lightness_female 0.7273 0.7262 0.999
60% lightness_male 0.6943 0.6943 1.000

Wall time: 229.2 min. Generated: 2026-04-17 17:09

Reference: Delhey K, Dale J, Valcu M, Kempenaers B (2019). Ecology Letters, 22(5): 726-736.